any of several small angular bones that in humans make up the wrist (carpus), and in horses, cows, and other quadrupeds the “knee” of the foreleg. They correspond to the tarsal bones of the rear or lower limb. Their number varies. Primitive vertebrates typically had 12. In modern amphibians, reptiles, and birds, the number is reduced by fusion. In humans there are eight, arranged in two rows. The bones in the row toward the forearm are the scaphoid, lunate, triangular, and pisiform. The row toward the fingers, or distal row, includes the trapezium (greater multangular), trapezoid (lesser multangular), capitate, and hamate. The distal row is firmly attached to the metacarpal bones of the hand. The proximal row articulates with the radius (of the forearm) and the articular disk (a fibrous structure between the carpals and malleolus of the ulna) to form the wrist joint.